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A noticeable increase in head width, resulting from the swelling of the whole body of the male during the aquatic phase, is unknown in families other than hynobiids that accomplish external fertilization and is related to male–male competition.

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This page is a summary of: Seasonal Changes in Body Shape and Mass in the Salamander, Hynobius nigrescens, Journal of Herpetology, June 1990, JSTOR,
DOI: 10.2307/1564217.
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