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  1. Tiny guardian of the spring: Description of the first Islamiinae (Mollusca, Hydrobiidae) from the Caucasus

    Article • ZooKeys, March 2026, Pensoft Publishers

    Prof. Assoc. Levan Mumladze

  2. Corrigendum to “Edaphobase 2.0: Advanced international data warehouse for collating and using soil biodiversity datasets” [Applied Soil Ecology 204 (2024) 105710]

    Article • Applied Soil Ecology, January 2026, Elsevier

    Prof. Assoc. Levan Mumladze, Professor Olaf Schmidt

  3. Open letter: A global call to strengthen national soil biodiversity action through coordination and harmonization

    Article • Plants People Planet, October 2025, Wiley

    Professor Olaf Schmidt, Dr Samuel Eziuzor, Prof. Assoc. Levan Mumladze, Dr Raza Ullah Khan

  4. Distribution, community composition and vulnerability of fish species in the Algeti River Basin (Georgia)

    Article • Caucasiana, June 2025, Pensoft Publishers

    Prof. Assoc. Levan Mumladze

  5. The complete mitochondrial genome of the widespread freshwater limpet Ferrissia californica

    Article • Mitochondrial DNA Part B, April 2025, Taylor & Francis

    Prof. Assoc. Levan Mumladze

  6. Insect Herbivory Releases More Nutrients in Warmer and Drier Forests

    Article • Global Biogeochemical Cycles, April 2025, American Geophysical Union (AGU)

    Prof. Assoc. Levan Mumladze

  7. The distribution of bat species across the underground sites of Georgia

    Article • Subterranean Biology, November 2024, Pensoft Publishers

    Prof. Assoc. Levan Mumladze

  8. New records of centipedes (Myriapoda: Chilopoda) in the fauna of Georgia, South Caucasus

    Article • Caucasiana, September 2024, Pensoft Publishers

    Prof. Assoc. Levan Mumladze

  9. A new Psammitis species (Araneae, Thomisidae) from an extinct volcano in Georgia with reevaluation of the generic position of Xysticus marmoratus Thorell, 1875

    Article • Caucasiana, April 2024, Pensoft Publishers

    Prof. Assoc. Levan Mumladze

  10. Checklist of digeneans (Platyhelminthes, Trematoda, Digenea) of Georgia

    Article • Biodiversity Data Journal, January 2024, Pensoft Publishers

    Prof. Assoc. Levan Mumladze

  11. New records and first DNA barcodes of water mites (Acari, Hydrachnidia) from Georgia

    Article • Ecologica Montenegrina, October 2023, Institute for Biodiversity and Ecology

    Prof. Assoc. Levan Mumladze

  12. Fish diversity assessed by eDNA detection methods in the Rioni River

    Article • Metabarcoding and Metagenomics, October 2023, Pensoft Publishers

    Prof. Assoc. Levan Mumladze

  13. Checklist of digeneans (Platyhelminthes, Trematoda, Digenea) of Georgia

    Article • August 2023, Pensoft Publishers

    Prof. Assoc. Levan Mumladze

  14. Species elevational richness gradient and species-area relationship in mountain vegetation of Javakheti highland (Georgia)

    Article • Caucasiana, July 2023, Pensoft Publishers

    Prof. Assoc. Levan Mumladze

  15. Phylogeographic patterns of genetic diversity in Pontastacus leptodactylus (Decapoda: Astacidae): is the hypothesis of the taxonomically rich genus Pontastacus true?

    Article • Zoological Journal of the Linnean Society, July 2023, Oxford University Press (OUP)

    Prof. Assoc. Levan Mumladze

  16. Bats of Georgia - an occurrence dataset from 1835 through 2022

    Article • Biodiversity Data Journal, June 2023, Pensoft Publishers

    Prof. Assoc. Levan Mumladze

  17. Ponticola alasanicus sp. n. (Gobiiformes, Gobiidae) from the Alazani River Basin, Georgia

    Article • Biodiversity Data Journal, May 2023, Pensoft Publishers

    Prof. Assoc. Levan Mumladze

  18. The first DNA-assisted record of Hemelytroblatta livida (Blattodea: Corydiidae) from Georgia with notes on Corydiinae species composition in the Caucasus

    Article • Caucasiana, May 2023, Pensoft Publishers

    Prof. Assoc. Levan Mumladze

  19. The record of Ametropus fragilis Albarda, 1878 (Ephemeroptera, Ametropodidae) from Georgia

    Article • Caucasiana, May 2023, Pensoft Publishers

    Prof. Assoc. Levan Mumladze

  20. Diatom endemism and taxonomic turnover: Assessment in high-altitude alpine lakes covering a large geographical range

    Article • The Science of The Total Environment, May 2023, Elsevier

    Prof. Assoc. Levan Mumladze

  21. The land snail family Clausiliidae (Gastropoda, Pulmonata, Stylommatophora) in Georgia: overview, novel records and a new species

    Article • Caucasiana, March 2023, Pensoft Publishers

    Prof. Assoc. Levan Mumladze

  22. Revealing the Stygobiotic and Crenobiotic Molluscan Diversity in the Caucasus: Part IV—Crenobiotic Belgrandiellinae Radoman, 1983 (Mollusca, Hydrobiidae) from Georgia

    Article • Diversity, March 2023, MDPI AG

    Prof. Assoc. Levan Mumladze

  23. Water bugs (Heteroptera: Gerromorpha, Nepomorpha) of the Caucasus ecoregion

    Article • The European Zoological Journal, February 2023, Taylor & Francis

    Prof. Assoc. Levan Mumladze

  24. The genus Acicula W. Hartmann, 1821 in the Caucasus (Gastropoda: Caenogastropoda: Aciculidae)

    Article • Zootaxa, February 2023, Magnolia Press

    Prof. Assoc. Levan Mumladze

  25. Annotated checklist of Georgian oribatid mites—II

    Article • Zootaxa, January 2023, Magnolia Press

    Prof. Assoc. Levan Mumladze

  26. Evidence of cryptic speciation in the Hygrobates calliger complex (Acariformes, Hydrachnidia, Hygrobatidae) with the description of two new species

    Article • Ecologica Montenegrina, December 2022, Institute for Biodiversity and Ecology

    Prof. Assoc. Levan Mumladze

  27. Development and application of a multilingual electronic decision-support tool for risk screening non-native terrestrial animals under current and future climate conditions

    Article • NeoBiota, October 2022, Pensoft Publishers

    Dr Marina Piria, Prof. Assoc. Levan Mumladze

  28. Risk of invasiveness of non-native fishes in the South Caucasus biodiversity and geopolitical hotspot

    Article • NeoBiota, October 2022, Pensoft Publishers

    Dr Marina Piria, Prof. Assoc. Levan Mumladze

  29. Species diversity and DNA barcode library of freshwater Molluscs of South Caucasus

    Article • Biodiversity Data Journal, September 2022, Pensoft Publishers

    Prof. Assoc. Levan Mumladze

  30. Species diversity and DNA barcode library of freshwater Molluscs of South Caucasus

    Article • April 2022, Pensoft Publishers

    Prof. Assoc. Levan Mumladze

  31. Check-list of chromosome numbers of the family Hygromiidae (Gastropoda: Stylommatophora) with new data on Circassina frutis

    Article • Travaux du Muséum National d’Histoire Naturelle “Grigore Antipa”, December 2021, Pensoft Publishers

    Prof. Assoc. Levan Mumladze

  32. A global agenda for advancing freshwater biodiversity research

    Article • Ecology Letters, December 2021, Wiley

    Dr Javier J Muzon, Prof. Assoc. Levan Mumladze, Dr Gregor Kalinkat

  33. Hydropower development in the Republic of Georgia and implications for freshwater biodiversity conservation

    Article • Biological Conservation, November 2021, Elsevier

    Prof. Assoc. Levan Mumladze

  34. A global-scale screening of non-native aquatic organisms to identify potentially invasive species under current and future climate conditions

    Article • The Science of The Total Environment, September 2021, Elsevier

    Dr Aurel Năstase, Prof. Assoc. Levan Mumladze

  35. Life histories of three Serrulinini species (Stylommatophora: Clausiliidae) from Georgia kept under laboratory conditions

    Article • Folia Malacologica, August 2021, Bogucki Wydawnictwo Naukowe

    Prof. Assoc. Levan Mumladze

  36. Oribatid (Acari: Oribatida) diversity in natural and altered open arid ecosystems of South-Eastern Caucasus

    Article • Pedobiologia, August 2021, Elsevier

    Prof. Assoc. Levan Mumladze

  37. The role of Anatolia in the origin of the Caucasus biodiversity hotspot illustrated by land snails in the genusOxychilus

    Article • Cladistics, July 2021, Wiley

    Prof. Assoc. Levan Mumladze

  38. Species diversity and distribution of freshwater molluscs of Javakheti Highlands (Republic of Georgia)

    Article • Biodiversity Data Journal, June 2021, Pensoft Publishers

    Prof. Assoc. Levan Mumladze

  39. Species diversity and distribution of freshwater molluscs of Javakheti Highlands (Republic of Georgia)

    Article • March 2021, Pensoft Publishers

    Prof. Assoc. Levan Mumladze

  40. Speaking their language – Development of a multilingual decision-support tool for communicating invasive species risks to decision makers and stakeholders

    Article • Environmental Modelling & Software, January 2021, Elsevier

    Dr Marina Piria, Prof. Assoc. Levan Mumladze

  41. The first unified inventory of non-native fishes of the South Caucasian countries, Armenia, Azerbaijan, and Georgia

    Article • Knowledge and Management of Aquatic Ecosystems, January 2021, EDP Sciences

    Prof. Assoc. Levan Mumladze

  42. Towards retrieving the Promethean treasure: a first molecular assessment of the freshwater fish diversity of Georgia

    Article • Biodiversity Data Journal, October 2020, Pensoft Publishers

    Prof. Assoc. Levan Mumladze

  43. Genetic diversity, phylogenetic position and morphometric analysis of Astacus colchicus (Decapoda, Astacidae): a new insight into Eastern European crayfish fauna

    Article • Integrative Zoology, October 2020, Wiley

    Prof. Assoc. Levan Mumladze

  44. Revealing the stygobiotic and crenobiotic molluscan biodiversity hotspot in Caucasus: Part I. The phylogeny of stygobiotic Sadlerianinae Szarowska, 2006 (Mollusca, Gastropoda, Hydrobiidae) from Georgia with descriptions of five new genera and twenty-on...

    Article • ZooKeys, August 2020, Pensoft Publishers

    Prof. Assoc. Levan Mumladze

  45. First survey of shallow-water Amphipoda along the Georgian Black Sea coast reveals new faunistic records and the unexpected Atlantic invader Melita nitida

    Article • Mediterranean Marine Science, June 2020, National Documentation Centre

    Prof. Assoc. Levan Mumladze

  46. Checklist of the freshwater fishes of Armenia, Azerbaijan and Georgia

    Article • Journal of Applied Ichthyology, May 2020, Hindawi Publishing Corporation

    Prof. Assoc. Levan Mumladze

  47. New remarkable record of Helicopsyche bacescui Orghidan and Botosaneanu, 1953 (Trichoptera, Helicopsychidae) from Georgia

    Article • Aquatic Insects, March 2020, Taylor & Francis

    Prof. Assoc. Levan Mumladze

  48. Faunal biodiversity research in the Republic of Georgia: a short review of trends, gaps, and needs in the Caucasus biodiversity hotspot

    Article • Biologia, December 2019, Springer Science + Business Media

    Prof. Assoc. Levan Mumladze

  49. Forest Restoration Potential in a Deforested Mountain Area: An Ecosociological Approach towards Sustainability

    Article • Forest Science, December 2019, Oxford University Press (OUP)

    Prof. Assoc. Levan Mumladze

  50. Materials to the fauna of aquatic bugs of the infraorder Nepomorpha (Heteroptera) of the Javakheti Highland, Georgia

    Article • rej, September 2019, KMK Scientific Press

    Prof. Assoc. Levan Mumladze

  51. Faunal Biodiversity Research in the Republic of Georgia: A Short Review of Trends, Gaps, and Needs in the Caucasus Biodiversity Hotspot

    Article • August 2019, MDPI AG

    Prof. Assoc. Levan Mumladze

  52. Redescription of Roncus crassipalpus Rafalski, 1949 (Pseudoscorpiones: Neobisiidae) from western Georgia

    Article • Arachnology, August 2019, Arachology

    Prof. Assoc. Levan Mumladze

  53. A new pseudoscorpion genus from western Georgia (Pseudoscorpiones: Neobisiidae: Cornuroncus n. gen.), with a key to all Neobisiinae genera

    Article • Zootaxa, June 2019, Magnolia Press

    Prof. Assoc. Levan Mumladze

  54. New freshwater invasive mollusc species in Georgia

    Article • Vestnik Zoologii, June 2019, De Gruyter

    Prof. Assoc. Levan Mumladze

  55. Phylogeny, phylogeography and hybridization of Caucasian barbels of the genus Barbus (Actinopterygii, Cyprinidae)

    Article • Molecular Phylogenetics and Evolution, June 2019, Elsevier

    Prof. Assoc. Levan Mumladze

  56. Parthenogenetic vs. sexual reproduction in oribatid mite communities

    Article • Ecology and Evolution, May 2019, Wiley

    Prof. Assoc. Levan Mumladze

  57. New species and new records of <i>Aphelinus</i> Dalman (Hymenoptera: Chalcidoidea: Aphelinidae) from Lagodekhi Reserve (Sakartvelo - Georgia), with diversity and distribution along an elevational gradient

    Article • TURKISH JOURNAL OF ZOOLOGY, March 2019, The Scientific and Technological Research Council of Turkey (TUBITAK-ULAKBIM) - DIGITAL COMMONS JOURNALS

    Prof. Assoc. Levan Mumladze

  58. Neobisium (Neobisium) moreoticum (Pseudoscorpiones: Neobisiidae) from Georgia

    Article • Arachnologische Mitteilungen Arachnology Letters, February 2019, Arachnologische Mitteilungen

    Prof. Assoc. Levan Mumladze

  59. Effect of ploughing and pesticide application on oribatid mite communities

    Article • International Journal of Acarology, February 2019, Taylor & Francis

    Prof. Assoc. Levan Mumladze

  60. Phylogeny, phylogeography and hybridization of Caucasian barbels of the genusBarbus(Actinopterygii, Cyprinidae)

    Article • November 2018, Cold Spring Harbor Laboratory Press

    Prof. Assoc. Levan Mumladze

  61. A New Cave Pseudoscorpion (Pseudoscorpiones: Neobisiidae) from Western Georgia

    Article • Arachnology, November 2018, Arachology

    Prof. Assoc. Levan Mumladze

  62. The first annotated checklist of mayflies (Ephemeroptera: Insecta) of Georgia with new distribution data and a new record for the country

    Article • TURKISH JOURNAL OF ZOOLOGY, March 2018, The Scientific and Technological Research Council of Turkey (TUBITAK-ULAKBIM) - DIGITAL COMMONS JOURNALS

    Prof. Assoc. Levan Mumladze

  63. Fish species composition, sex ratio and growth parameters in Saghamo Lake (Southern Georgia)

    Article • Biologia, January 2018, Springer Science + Business Media

    Prof. Assoc. Levan Mumladze

  64. Patterns of oribatid mite species diversity: testing the effects of elevation, area and sampling effort

    Article • Experimental and Applied Acarology, July 2017, Springer Science + Business Media

    Prof. Assoc. Levan Mumladze

  65. List of leaf beetles (Coleoptera: Chrysomelidae) from Lagodekhi reserve with new records for Transcaucasia and Georgia

    Article • Zootaxa, June 2017, Magnolia Press

    Prof. Assoc. Levan Mumladze

  66. Beyond elevation: testing the climatic variability hypothesis vs. Rapoport’s rule in vascular plant and snail species in the Caucasus

    Article • Biological Journal of the Linnean Society, April 2017, Oxford University Press (OUP)

    Prof. Assoc. Levan Mumladze

  67. An inverse elevational species richness gradient of Caucasian vascular plants and Encyrtidae (Hymenoptera, Chalcidoidea)

    Article • Ecoscience, April 2017, Taylor & Francis

    Prof. Assoc. Levan Mumladze

  68. Annotated checklist of Georgian oribatid mites 

    Article • Zootaxa, March 2016, Magnolia Press

    Prof. Assoc. Levan Mumladze

  69. Article • Zootaxa, March 2016, Magnolia Press

    Prof. Assoc. Levan Mumladze

  70. Oribatid mite communities along an elevational gradient in Sairme gorge (Caucasus)

    Article • Experimental and Applied Acarology, March 2015, Springer Science + Business Media

    Prof. Assoc. Levan Mumladze

  71. Sympatry without co-occurrence: exploring the pattern of distribution of two Helix species in Georgia using an ecological niche modelling approach

    Article • Journal of Molluscan Studies, June 2014, Oxford University Press (OUP)

    Prof. Assoc. Levan Mumladze

  72. Endemic land molluscs in Georgia (Caucasus): how well are they protected by existing reserves and national parks?

    Article • Journal of Molluscan Studies, February 2014, Oxford University Press (OUP)

    Prof. Assoc. Levan Mumladze

  73. Invasive Carassius carp in Georgia: Current state of knowledge and future perspectives

    Article • Current Zoology, December 2013, Oxford University Press (OUP)

    Prof. Assoc. Levan Mumladze

  74. Systematics and Evolutionary History of Large Endemic Snails from the Caucasus (Helix buchiiandH. goderdziana) (Helicidae)

    Article • American Malacological Bulletin, August 2013, American Malacological Society

    Prof. Assoc. Levan Mumladze

  75. Oribatid mite colonization of sand and manganese tailing sites

    Article • Acarologia, June 2013, Les Amis d'Acarologia

    Prof. Assoc. Levan Mumladze

  76. Compositional patterns in Holarctic peat bog inhabiting oribatid mite (Acari: Oribatida) communities

    Article • Pedobiologia, January 2013, Elsevier

    Prof. Assoc. Levan Mumladze

  77. Shell size differences in Helix lucorum Linnaeus, 1758 (Mollusca: Gastropoda) between natural and urban environments

    Article • TURKISH JOURNAL OF ZOOLOGY, January 2013, The Scientific and Technological Research Council of Turkey (TUBITAK-ULAKBIM) - DIGITAL COMMONS JOURNALS

    Prof. Assoc. Levan Mumladze

  78. <p><strong>Landscape distribution of oribatid mites (Acari, Oribatida) in Kolkheti National Park (Georgia, Caucasus)*</strong></p>

    Article • Zoosymposia, December 2011, Magnolia Press

    Prof. Assoc. Levan Mumladze

  79. Palaeoclimatic models help to understand current distribution of Caucasian forest species

    Article • Biological Journal of the Linnean Society, November 2011, Oxford University Press (OUP)

    Prof. Assoc. Levan Mumladze

  80. Forest snail faunas from Georgian Transcaucasia: patterns of diversity in a Pleistocene refugium

    Article • Biological Journal of the Linnean Society, January 2011, Oxford University Press (OUP)

    Prof. Assoc. Levan Mumladze

  81. Comparison of Earthworms (Lumbricidae) and Oribatid Mite (Acari, Oribatida) Communities in Natural and Urban Ecosystems

    Article • Vestnik Zoologii, January 2011, National Academy of Sciences of Ukraine (Co. LTD Ukrinformnauka) (Publications)

    Prof. Assoc. Levan Mumladze

  82. Unisexual rock lizard might be outcompeting its bisexual progenitors in the Caucasus

    Article • Biological Journal of the Linnean Society, September 2010, Oxford University Press (OUP)

    Prof. Assoc. Levan Mumladze

  83. Botanical and zoological remains from an early medieval grave at Tsitsamuri, Georgia

    Article • Vegetation History and Archaeobotany, October 2008, Springer Science + Business Media

    Prof. Assoc. Levan Mumladze